Know Your Natives – Rattle Weed

Rattle weed (Astragalus canadensis) of the Pea (Fabaceae) family, occurs throughout the US except for Arizona, Florida and the far Northeast.  It is also found throughout Canada.  In Arkansas, rattle weed occurs in scattered counties across the state, with most occurrences in the Interior Highlands of the northwestern half of the state.  Also known as Canadian milk vetch, rattle weed is an herbaceous perennial.  It is found in sunny and wet to somewhat dry prairies, open woodlands, roadsides, thickets and stream banks.

The plant has shallow, widely branched near-surface roots.  Spring growth sprouts at numerous points along the roots.  Early leaves develop from erect, light green stems that eventually grow to 1½ – 4 feet tall.  Plants in strong sun may develop reddish stems.  The plant’s long slender stems, which have similar girth from ground to inflorescence, may recline later in the growing season.  The stems are hollow and somewhat square in cross-section, with minor ridging all around.  Secondary stems arise from leaf axils, and leaf scars occur all around the stems.  The stems may be smooth or pubescent.

Rattle weed - Astragalus canadensisPhoto 1:  New stems of rattle weed emerge in early spring from various points along branched roots.  With age, colonies may form.

Medium-green, odd-pinnate compound leaves may be a foot long with up to about 31 widely spaced leaflets.  Opposite (mostly) leaflets decrease in size towards leaf tips.  Insignificant stipules are triangular and pointed.  The rachis is grooved along its upper length.  The smoothed-edged, oblong to elliptic leaflets on short petiolules are up to about 1½ inches long and ¾ inch wide, with rounded bases and rounded to emarginate tips.  Leaflets are generally smooth or slightly hairy on upper surfaces, with stiff short hairs on lower surfaces.

Rattle weed - Astragalus canadensisPhoto 2:  Ten-inch mid-stem leaves of rattle weed in late spring.  Leaflets are normally opposite, but may occasionally be alternate (see upper leaf).

The inflorescence of rattle weed, occurring in late spring, consists of multiple upright racemes of densely clustered white to pale greenish-yellow, pea-shaped flowers which grow from axils of upper leaves along primary and secondary stems.  Racemes, on long stems (peduncles), have an elongated cone shape with 50 to 120 or more flowers whorled evenly all around the upper two-thirds of the stem.  Each flower bud is subtended by a bract which withers as the flower opens.  Raceme and stem combined are up to 7 inches long.

Rattle weed - Astragalus canadensisPhoto 3:  Raceme with flower buds inclined upward.  Each bud is subtended by a bract.

Flowers of rattle weed open in succession from the bases of the racemes to tops.  Open flowers toward the bottom of the racemes tilt downward while higher flowers are horizontal and then tilted upward.   Flowers, ½ to ¾ inch long with a greenish-white, hairy calyx with five triangular pointed lobes, have a tubular corolla composed of five creamy greenish-white petals.  The flowers are pea-like and elongated with a petal forming the upper banner, two lateral petals and with two petals forming the keel.  The stamens, pistil and style are enclosed by the keel.

Rattle weed - Astragalus canadensisPhoto 4:  Rattle weed racemes grow from multiple leaf axils near tops of long, slender stems.

Flowers are replaced by inflated, smooth and oval green pods with long pointed tips.  The pods, about ½ to ¾ inch long, become dark brown and woody in texture with maturity.  The dozen or so seeds in a dry pod become loose and rattle when stems are shaken, thus the common name “rattle weed.”  The pods are persistent.

Rattle weed - Astragalus canadensisPhoto 5:  Maturing seed pods.  All flowers do not produce seed.

Rattle weed, as a fairly tall specimen plant, can be a good garden plant.  Its compound leaves add textural variety and multiple attractive flower clusters are born at the top of the plant.  It is not known to be aggressive either from seed or root.  The nectar is used by various butterflies and other insects.  Seeds are eaten by song birds and turkey.  The foliage may be eaten by deer and other herbivores.

Article and photographs by ANPS member Sid Vogelpohl

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Know Your Natives – Jack-in-the-Pulpit & Green-Dragon

Jack-in-the-pulpit and green-dragon, in the Arum (Araceae) Family, are herbaceous perennial monocots that occur throughout the state in similar habitats.  Both are smooth overall and hairless.  Plants range from 2 inches to 2 feet in height.  Habitats include mesic (well-balanced moisture) deciduous woodlands and thickets and hillside seeps with light shade and humus-rich soil.  Both are dioecious; that is, the flowers are unisexual, staminate or pistillate, such that two plants are required for cross-pollination and fertilization. In general a plant will be staminate for several years, contributing only pollen to the reproductive process, until enough energy has accrued in the corm to produce seeds and fruits. The sex change may be brought about by age or health of the plant.  Corms, about an inch below the surface, may grow to 1½ inches across with a few fibrous roots.  Both species produce an elongated cluster (corn-cob-like) of bright shiny red berries that appear identical for both plants (photo 1).  One- or two-year-old plants of both species have a single leaf with three leaflets and appear identical.  Both species produce clonal plantlets, and thick-standing colonies of varying sized plants can develop.  Leaves and flowers unfurl initially from a pointed sheaf arising from each corm.  Inflorescences, on top of stout fleshy upright stalks, emerge at the same time as leaves.

Green Dragon - Arisaema dracontiumPhoto 1:  Mature cluster of green-dragon berries.  Cluster on Jack-in-the pulpit would appear identical.

Berries (¼ inch across) enclosing one to several seeds, at first green, become bright red as female plants begin to wither in late summer and fall.  Berries remain attached to the dry spadix, resulting in an ovoid mass of showy berries up to 2 inches long which stay on the stem even when stem has dried.  A cluster can consist of up to 150 berries, each with 1 to 3 rounded flat-sided light tan seeds.

Either plant is an excellent woodland garden plant, being easy to cultivate and requiring little care.  Along with the interesting characteristics of the vegetation, the cluster of berries in late summer and fall is attractive.  Birds and mammals eat the berries, but berries of both plants should be considered poisonous to people.

Jack-in-the-Pulpit

Jack-in-the-pulpit (Arisaema triphyllum [formerly Arisaema atrorubens]) is also known as Indian-turnip because cooked corms were eaten by Native Americans.  A mature corm produces one or two large glossy compound leaves on stout fleshy stalks.  Typically, three smooth-edged leaflets emanate from a common point at the top of each leaf stem (photo 2), though leaves can occasionally have five leaflets (photo 3).  Leaflets, up to a foot long and up to 8 inches wide, are broadly oval to elliptic with tapering points.  Leaflets of both plants have pinnate venation which stops near leaf edges.

 Jack-in-the-Pulpit - Arisaema triphyllumPhoto 2:  Characteristic shape of Jack-in-the-pulpit’s compound leaves.  Typical plants have only three leaflets.  Note venation.

Jack-in-the-Pulpit - Arisaema triphyllumPhoto 3:  Occasional Jack-in-the-pulpit plants have leaves with five leaflets each.

The inflorescence, initially wrapped by stipules at the base of leaf or leaves, has a spathe (widened hood or pulpit) rising above the central protruding spadix (Jack) (photo 4).  Exterior of the spathe is usually green or purple and the inside usually striped greenish white (along veins that parallel flower stalk) and purple (between veins).  The lower portion of the spathe is a tube-like sheath wherein numerous tiny male and/or female flowers, tightly bound to the spadix, are hidden.  Structures on the spadix standing above male and female flowers are infertile.  Pollinating flies are trapped in the lower portion of the spathe (photo 5).

Jack-in-the-Pulpit - Arisaema triphyllumPhoto 4:  Inflorescence of Jack-in-the-Pulpit showing over-arching spathe and post-like spadix.

Jack-in-the-Pulpit - Arisaema triphyllumPhoto 5:  Developing berries of Jack-in-the-pulpit.  The spathe has begun to dry.  Insect carcasses litter the base of the spathe.

Green-Dragon

Green-dragon or dragon-root (Arisaema dracontium) is very similar to Jack-in-the pulpit, with main differences being in leaves and inflorescence.  Green-dragon usually has only one large compound leaf with 7 to 15 lance-shaped leaflets (photo 6).  Green-dragon has a greenish spathe that is less prominent and the spadix is considerably longer, thinner, and tapered, extending upward (the dragon’s tongue) around the top of the spathe (photo 7).  Mature green-dragon leaves have the central and largest leaflet unfurling first.  At the base of the central leaflet, other leaflets branch off, forming a semi-circle parallel to the ground [the dragon’s wings?] (photo 8).

Green Dragon - Arisaema dracontiumPhoto 6:  Leaves of green-dragon.  The largest central leaflet ties directly to the leaf stalk while other leaflets grow from a “branch” on either side. Note venation.

Arisaema dracontiumPhoto 7: Green-dragon inflorescence.  Spadix is several times longer than spathe. (Plant seen behind green-dragon is May-apple.)

Green Dragon - Arisaema dracontiumPhoto 8:  A new leaf and flower of green-dragon.  Leaf may suggest dragon’s wings.  (Plant seen behind green-dragon is celandine poppy.)

Article and photographs by ANPS member Sid Vogelpohl

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Neat Plant Alert – Scarlet Beardtongue

The recent ANPS field trip to Poison Springs Natural Area and nearby back roads was very productive! Trip leader Meredith York took us to see some great plants, many of which are rare in Arkansas.

The most spectacular sight of the day was without a doubt Scarlet Beardtongue (Penstemon murrayanus).

Scarlet Beardtongue - Penstemon murrayanus

Scarlet Beardtongue photographed along County Road 423 in Nevada County

Found only in Nevada and Ouachita counties in southwest Arkansas, Scarlet Beardtongue is more common to the southwest in the eastern half of Texas.

Scarlet Beardtongue - Penstemon murrayanus

Our largest Beardtongue – the blooming stalks can reach over 6 feet in height by the end of the bloom season

Scarlet Beardtongue - Penstemon murrayanus

The bright red flowers are a favorite of Ruby-throated hummingbirds

Article and photographs by ANPS member Eric Hunt

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Know Your Natives – Goat’s Rue

Goat’s-rue (Tephrosia virginiana) of the Pea (Fabaceae) Family occurs throughout the eastern U.S. from Texas and Minnesota to the Atlantic, except for Vermont and Maine.  In Arkansas, it occurs mostly statewide except for counties along the Mississippi River.  Other common names for goat’s-rue include Virginia tephrosia, catgut and devil’s-shoestrings.  The genus name, Tephrosia, is based on the Greek word tephros meaning “ash-colored” or “hoary” due to the white or grayish hairs covering the plant.  The name “goat’s-rue” is based on the plant having been fed to goats with the belief that it increased milk production, but that use has been discontinued.  The name “devil’s-shoestrings” relates to the plant’s long, tough fibrous roots.  This plant, as does many other plants in the Pea Family, has a symbiotic relationship with a soil bacterium (rhizobium) whereby the plant converts (fixes) atmospheric nitrogen to a usable form within root nodules.

Goat’s-rue is an herbaceous perennial legume favoring sunny sites with sandy to rocky, dry soils of glades, open woodlands and prairies, generally on acidic soils.  Plants consist of sparingly branched stems bearing odd-pinnate compound leaves and that terminate with an inflorescence of pea-like flowers.  Stems, from a caudex, can be several or numerous.  Early growth of plants have an overall light green color which changes to grayish green with maturity due to plant hairs.  Hairiness varies from a fine, soft pubescence on upper leaf surfaces to densely hairy on stems and seed pods.  Plants, up to two feet tall, tend to recline later in the growing season.

Goat's Rue - Tephrosia virginianaPhoto 1:  Early spring growth of goat’s-rue in a rocky and sunny glade.

Compound alternate leaves are odd-pinnate, having 14 to 24 lateral leaflets and one terminal leaflet.  Leaflets, up to an inch long and ¼ inch wide, are oblong to narrowly elliptic with entire margins and gradually terminate at a sharp tip.  Lateral leaflets tend to be opposite, but may be offset across the rachis (main axis of leaf).   Leaves have short petioles (attachment of rachis to stem) and leaflets have short petiolules (attachment of leaflet to rachis).  The upper surface of the rachis has a central grove.  Central veins of leaflets are prominent on the under sides.

Goat’s-rue blooms in late April or May, with the inflorescence consisting of dense racemes several inches long at the tops of the stems.  Five light green sepals are fused to form a calyx with five teeth.  The calyx is densely hairy on the outside, as are the pedicels and stems.  Buds and flowers face in all directions around the inflorescence.  Bicolored, pea-like flowers, ¾ inch long and across, have a broad upright petal (banner or standard) in shades of cream and yellow and a pair of lateral, forward projecting petals (wings) that are deep rosy pink.  The wings enclose a pair of smaller petals fused at their apex to form a keel which encloses ten stamens and a pistil.

Goat's Rue - Tephrosia virginianaPhoto 2:  Close-up of goat’s-rue inflorescence showing hairiness of plant and bicolored flowers.

Goat's Rue - Tephrosia virginianaPhoto 3:  Goat’s-rue plant at peak bloom.

Fruit, which form in mid-summer, are flattened, 2-inch long hirsute pods situated on the stems in widely spreading fashion.  They are initially light green, but later turn brown with maturity.  The pods contain flattened, kidney-shaped mini-beans.  When mature and dry, the pods split, with the halves twisting in a cork-screw fashion, ejecting seeds a short distance.

Goat's Rue - Tephrosia virginianaPhoto 4:  Densely pubescent seed pods of goat’s-rue firmly fixed in various directions from stem.

Goat’s-rue is a beautiful native plant that looks attractive in a natural setting or a sunny garden with well-drained soil.  Nectar, pollen and leaves attract leaf-cutting bees and other insects.  However, plants may be difficult to establish from seed or pots, and they are extremely difficult to transplant due to their large roots.  The plant contains rotenone (used as an insecticide and to kill invasive fish) and should be considered toxic to people and many animals.

A second member of this genus also occurs in Arkansas: hoary-pea (Tephrosia onobrychoides).  It is found in prairies and fields of the West Gulf Coastal Plain and Arkansas Valley.  Flowers of hoary-pea are smaller and occur widely spaced in racemes on tall stems held well above the leaves.  Leaves and leaflets, too, are more widely spaced than in goat’s-rue.  The solid-colored flowers of this species start out as pale pink to almost white when flowers first open but then change to a deep rose-red as the flowers fade.

Article and photographs by ANPS member Sid Vogelpohl

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Neat Plant Alert – Copper Iris

Today I pulled out my waterproof boots and went in search of Copper Iris (Iris fulva) in Little Rock. Mid-April is bloom season for Copper Iris, and I knew there was a population in the Fouche Creek bottomlands owned by the City of Little Rock in the area south of the I-30/440/530.

Lucky for me it was in bloom just a few dozen yards off the access road!

Iris fulva

Copper iris growing in a seasonally wet bottomland forest in south Little Rock

The species is more common in counties closer to the Mississippi River and can be sometimes spotted growing in ditches along backroads in those counties.

Article and photographs by ANPS member Eric Hunt

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Know Your Natives – Wood-betony

One of our more showy spring wildflowers is wood-betony (Pedicularis canadensis) in the Broomrape (Orobanchaceae) family.

Wood-betony can be found across much of eastern North America with some isolated populations in Colorado and New Mexico. In Arkansas it is found in most counties except those directly along the Mississippi River.

The foliage is hairy and lobed, almost resembling a fern but thicker, especially in early spring before the inflorescence emerges. It can sometimes have an attractive red tint to it.

Wood-betony (Pedicularis canadensis)

Photographed in Boyle Park, Little Rock

Flowers are produced at the top of a short inflorescence. They are usually clear yellow but some populations are bicolored purple/white or yellow/white.

Wood-betony (Pedicularis canadensis)

Photographed along the Lost Valley Trail, Buffalo National River, Newton County

Wood-betony (Pedicularis canadensis)

Photographed along the Lost Valley Trail, Buffalo National River, Newton County

Wood-betony grows in a variety of habitats, generally preferring woodlands, prairies, savannahs and riparian areas. It can become established in the artificial ‘prairies’ created next to roads where it takes advantage of abundant water runoff and no competition for light.

Wood-betony (Pedicularis canadensis)

Photographed along Highway 9 in the Ouachita National Forest, Perry County

Long-tongued bees such as bumblebees and mason bees are a major visitor to the flowers.

Wood-betony (Pedicularis canadensis)

Photographed along Highway 9 in the Ouachita National Forest, Perry County

Article and photographs by ANPS member Eric Hunt

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Native Plant Report – Ouachita National Trail – Perry County

Spring is in full swing in the Ouachita Mountains! The first ANPS Native Plant hike of 2015 was on April 11. Seven enthusiastic participants enjoyed a spectacular spring day in the forest.

A full hike report will appear in the Fall 2015 Claytonia. Today’s post is a quick visual taste of what was seen along the Ouachita National Trail in Perry County.

Hike participants were leaders Virginia McDaniel & Eric Hunt along with Don Ford, Becky Hardin, Mary McDaniel, Lynna Schoenert, and John Simpson.

Perfoliate bellwort - Uvularia perfoliata

Perfoliate bellwort – Uvularia perfoliata

Iris cristata

Dwarf Crested Iris – Iris cristata

Halesia carolina

Silverbells – Halesia carolina

Rhododendron prinophyllum

Early Azalea – Rhododendron prinophyllum

Osmundastrum cinnamomeum

Cinnamon fern – Osmundastrum cinnamomeum

Hamamelistes spinosus

Spiny Witch Hazel Gall Aphid – Hamamelistes spinosus – Growing on Common Witch Hazel – Hamamelis virginiana

Article and photographs by ANPS member Eric Hunt

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Know Your Natives – Wild Hyacinth

Wild hyacinth (Camassia scilloides) of the Agavaceae (Century Plant) family, formerly of the Liliaceae (Lily) Family, is also known as Atlantic camas and eastern camas.  It is found in the U.S. from Texas to Wisconsin and eastward to Georgia and Pennsylvania, with scattered occurrences in the Carolinas and Virginias.  In Arkansas it is found primarily in the highlands of the northwestern two-thirds of the state, with additional occurrences in the Grand Prairie and the western Gulf Coastal Plain.  Favorable sites for wild hyacinth have light shade to full sun, with moist, rich, loamy soil along stream terraces, in bottomland forests, glades and prairies.

The genus name is a combination of Native American words “camas” and “quamash” for “sweet” in reference to its bulb which was an important food source for Native Americans.*  The epithet “scilloides” is from Latin for “like Scilla” for a European plant with similar bulbs and leaves.  Wild hyacinth has spikes of flowers somewhat similar to the introduced garden hyacinth (Hyacinthus orientalis).

Wild hyacinth, a long-lived perennial, consists of a rosette of long, arching to reclined leaves and a flower stalk 1½-2 feet tall growing from a bulb with fibrous roots.  All parts of this ephemeral plant are glabrous (hairless).  Leaves, less than one inch wide and 6-12 inches long, are medium to dark green with smooth margins.  They have raised midribs on the undersides and are folded lengthwise in the lower halves.  They are generally somewhat floppy and twisted.

An erect, slender, light to medium-green, 1-2 foot tall stalk terminates in a raceme of flowers along the upper third.  The stalk is leafless except for several sterile bracts that may occur just below the raceme.  Each flower initially grows from the axil of a bract, with most of the bracts in the inflorescence falling off as the flowers open.  The bracts, linear and up to ¾ inch long, are the same color as the stem.

1 - Camassia scilloidesPhoto 1:  Wild hyacinth inflorescence stalk emerging in spring from rosette of leaves.  The lower portion of leaves are often folded and twisted.  The flowers occur in axils of linear bracts which are later deciduous.

2- Camassia scilloidesPhoto 2:  In wild hyacinth, most of the bracts in the inflorescence have fallen by the time the flowers open.  In this photo, only a few bracts below the raceme remain.

Flowering, in late spring, lasts about two weeks, with individual flowers lasting two to three days and opening successively from bottom to the top.  Flowers, 3/4 to 1 inch across, each consist of three sepals, three petals, six stamens with bright yellow anthers, and a light yellow, ovoid ovary topped by a long, slender style.  The light blue-violet to nearly white oblong tepals (sepals and petals) are widely flared.  Slender anthers and styles are about the same diameter and length.  Flowers, pollinated by bees, have a pleasant scent.

3 - Camassia scilloidesPhoto 3:  Wild hyacinth has relatively large ovaries with long slender styles.  Note two possible pollinator insects can be seen in this photo.

4 - Camassia scilloidesPhoto 4:  Tiger Swallowtail nectaring on wild hyacinth.  Note the lower flowers fading while upper flowers are still in bud or just opening.

Fertilized flowers are replaced by a three-loculed (chambered) capsule 1/3 inch long and wide.  Locules appear somewhat inflated, with dark green lines demarking the locule junctions.  Relatively large, shiny black seeds are limited to a dozen or so per capsule.  Wild hyacinth reproduces readily by seed.

5 - Camassia scilloidesPhoto 5:  Seed capsules of wild hyacinth become brown as they mature and as the plant begins to goes dormant.

Wild hyacinth is a great ephemeral to have in a garden, especially in its own space where its character can be fully appreciated.  Its appearance in late winter to early spring is a welcome sight.  The flowers attract bees, flies, butterflies and wasps.

Footnote:  Prairie wild hyacinth (Camassia angusta), also called prairie camas, is known to occur in Arkansas in Benton, Washington, Sebastian and Logan Counties.  Prairie wild hyacinth can be differentiated from wild hyacinth in that its flowers bloom several weeks later and are slightly smaller, the bracts below the inflorescence are more numerous and persistent, and the capsules are longer than wide.

*Bulbs of wild hyacinth were a highly nutritional food for Native Americans and early explorers.  Bulbs were used raw, baked, roasted, boiled or dried.  Remains of wild hyacinth (and other plant species) have been identified in earth ovens (middens) used by Native Americans.  For a detailed article relating to middens and plant remains studied at Camp Bowie, Texas, see: www.texasbeyondhistory.net/bowie/middentell.html.  CAUTION:  wild hyacinth bulbs and leaves look similar to those of the white-flowered death camas (Toxicoscordion nuttallii), a highly poisonous native plant.

Article and photographs by ANPS member Sid Vogelpohl

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Know Your Natives – Spring Spider-Lily

Spring Spider-lily

One of the most spectacular sights along the roads of central and southern Arkansas is the Spring Spider-lily (Hymenocallis liriosme).

Hymenocallis liriosme - Spring Spider Lily

A spectacular mass-blooming along US Highway 63 in the Grand Prairie, between Hazen and Stuttgart, Arkansas

Hymenocallis is a member of the Amaryllis family (Amaryllidaceae), and ranges from southern North America through the Caribbean and Central America and northern South America.

North America is home to 19 species of Spider-lily with 2 being found in Arkansas.

In Arkansas the Spring Spider-lily is found along the central Arkansas River Valley, in the Grand Prairie of southeastern Arkansas and in the Coastal Plain along the border with Louisiana. It is absent from the Ouachita and Ozark mountains.

Outside of Arkansas it is found in Louisiana, eastern Texas, southeastern Oklahoma, and is rare is Mississippi and Alabama.

Hymenocallis liriosme - Spring Spider Lily

The flowers have an elegant yellow-green eye in the center and are extremely fragrant

For most of the first half of the 20th century, Spring Spider-lily was considered a variety of the Northern Spider-lily (Hymenocallis occidentalis), which also occurs in Arkansas. In 1951 taxonomists recognized that the two species were in fact different and Spring Spider-lily was given species status.

Hymenocallis liriosme - Spring Spider Lily

Growing in standing water in the Grand Prairie between Hazen and Stuttgart along US Highway 63

It is very easy to distinguish Spring Spider-lily from Northern Spider-lily. Spring Spider-lily grows in large colonies in wet habitats, often in standing water, and blooms in mid-spring. The flowers have a prominent yellow-green eye in the center of the flower.

Northern Spider-lily grows in drier, upland habitats as single plants or small colonies. It blooms in mid-summer and the flowers have a much smaller yellow-green eye.

Spring Spider-lily is easily grown in the garden and makes an excellent rain garden or wetland garden specimen. It is available from commercial nurseries and is frequently shared as a pass-along plant in gardening circles.

Article and photographs by ANPS member Eric Hunt

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Native Plant Report – Warren Prairie

These photos were taken last week at Warren Prairie Natural Area.

Of special interest is the tiny Geocarpon minimum, one of the world’s rarest plants, and the slightly larger carnivorous sundew plant (watch your toes!).

This area will be featured in field trips and programs at the spring meeting of the Arkansas Native Plant Society, April 17-19, 2015.

Interested? Click here for full details.

Geocarpon minimum Warren Prairie

Geocarpon (Geocarpon minimum) Warren Prairie

Geocarpon minimum Warren Prairie

Geocarpon (Geocarpon minimum) at Warren Prairie

Sundew (Drosera brevifolia) Warren Prairie

Sundew (Drosera brevifolia) at Warren Prairie

Anemone caroliniana Warren Prairie

Carolina anemone (Anemone caroliniana) at Warren Prairie

Anemone caroliniana Warren Prairie

Carolina anemone (Anemone caroliniana) at Warren Prairie

Texas Saxifrage (Saxifraga texana) Warren Prairie

Texas Saxifrage (Micranthes texana) at Warren Prairie

Article and photographs by ANPS member Michael Weatherford

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